I used to run Nitto Dune Grapplers, tho I really didn't care much for the look of the treads. That said, they were excellent in the sand. They were pretty much great for dry pavement and sand, which is this country's environment 99% of the time. They performed average to bad on anything else. (Avg in wet roads, bad in mud). Nitto stopped making the Dune Grapplers in most sizes, including ours so I went with the KO2s.
Nearly all locals in their Land Cruisers run on dedicated sand tires out here, which are technically illegal on the road. They often call them "balloon tires". They are very rounded and meant to be aired down significantly.. as in single digit pressures! They have a high nylon / polyeurethane content.
This is the best pic of the balloon tires the locals use:
Here's a somewhat blurry depiction of what is happening when you air down your tire pressures:
As you can see, you gain a ton of contact distributing pressure giving more "float" to stay on top of the sand. The same technique is useful on rock crawling, but not quite as low pressures. Here's a quick guide on recommended pressures: